The dog ( Canis lupus familiaris ,pronounced /ˈkeɪnɨs ˈluːpəs fəˈmɪli.ɛərɨs/ ) is a domesticated form of the wolf, a member of the Canidae family of the order Carnivora. The term is used for both feral and pet varieties. The domestic dog has been one of the most widely kept working and companion animals in human history. The word "dog" may also mean the male of a canine species, as opposed to the word "bitch" for the female of the species.
The dog quickly became ubiquitous across culture across the world, and was extremely valuable to early human settlements. For instance, it is believed that the successful emigration across the Bering Strait might not have been possible without sled dogs. Dogs perform many roles for people, such as hunting, herding, protection, assisting police and military, companionship, and, more recently, aiding handicapped individuals. This versatility, more than almost any other known animal, has given them the nickname "Man's best friend" in the western world. Currently, there are estimated to be 400 million dogs in the world.
Over the 15,000 year span that the dog had been domesticated, it diverged into only a handful of landraces, groups of similar animals whose morphology and behavior have been shaped by environmental factors and functional roles. As the modern understanding of genetics developed, humans began to intentionally breed dogs for a wide range of specific traits. Through this process, the dog has developed into hundreds of varied breeds, and shows more behavioral and morphological variation than any other land mammal. For example, height measured to the withers ranges from a few inches in the Chihuahua to a few feet in the Irish Wolfhound; color varies from white through grays (usually called "blue'") to black, and browns from light (tan) to dark ("red" or "chocolate") in a wide variation of patterns; coats can be short or long, coarse-haired to wool-like, straight, curly, or smooth. It is common for most breeds to shed this coat.
Etymology and related terminology
Dog is the common use term that refers to members of the subspecies Canis lupus familiaris . The term is sometimes used to refer to a wider range of species: it can be used to refer to some belonging to the family Canidae, which includes foxes, jackals, Bush Dog, the cape dog Lycaon and coyotes and many others; or it can be used to refer to the subfamily of Caninae, or the genus Canis, also often called the "true dogs," which genus includes only the wolf, jackal, coyote, and dog. Some members of the family have "dog" in their common names, such as the Raccoon Dog and the African Wild Dog. A few animals have "dog" in their common names but are not canids, such as the prairie dog and the dog fish.
The English word dog comes from Middle English dogge , from Old English docga , a "powerful dog breed". The term may derive from Proto-Germanic *dukkōn , represented in Old English finger-docce ("finger-muscle"). The word also shows the familiar petname diminutive -ga also seen in frogga "frog", picga "pig", stagga "stag", wicga "beetle, worm", among others. Due to the archaic structure of the word, the term dog may ultimately derive from the earliest layer of Proto-Indo-European vocabulary, reflecting the role of the dog as the earliest domesticated animal.
Hound was traditionally the general word for all domestic canines, and dog was reserved for mastiffs and similar breeds. By the 16th century, dog had become the general word, and hound had begun to refer only to types used for hunting. Hound , cognate to German Hund , Dutch hond , common Scandinavian hund , and Icelandic hundur , is ultimately derived from the Proto-Indo-European *kwon- "dog", found in Welsh ci (plural cwn ), Latin canis , Greek kýōn , Lithuanian šuõ .
In breeding circles, a male canine is referred to as a dog , while a female is called a bitch (Middle English bicche, from Old English bicce, ultimately from Old Norse bikkja). A group of offspring is a litter . The father of a litter is called the sire , and the mother is called the dam . Offspring are generally called pups or puppies , from French poupée , until they are about a year old. The process of birth is whelping , from the Old English word hwelp , (cf. German Welpe, Dutch welp, Swedish valp, Icelandic hvelpur) .
Taxonomy
The domestic dog was originally classified as Canis familiaris and Canis familiarus domesticus by Carolus Linnaeus in 1758, and was reclassified in 1993 as Canis lupus familiaris , a subspecies of the gray wolf Canis lupus , by the Smithsonian Institution and the American Society of Mammalogists. Overwhelming evidence from behavior, vocalizations, morphology, and molecular biology led to the contemporary scientific understanding that a single species, the gray wolf, is the common ancestor for all breeds of domestic dogs ; however, the timeframe and mechanisms by which dogs diverged are controversial.
History and evolution
Main article: Origin of the domestic dogDomestic dogs inherited a complex social hierarchy and behaviors from their wolf ancestors. Dogs are pack animals with a complex set of behaviors related to determining each dog's position in the social hierarchy, and they exhibit various postures and other means of nonverbal communication that reveal their states of mind. These sophisticated forms of social cognition and communication may account for their trainability, playfulness, and ability to fit into human households and social situations, and these attributes have earned dogs a unique relationship with humans despite being potentially dangerous apex predators.
Although experts largely disagree over the details of dog domestication, it is agreed that human interaction played a significant role in shaping the subspecies. Shortly after domestication, dogs became ubiquitous in human populations, and spread throughout the world. Emigrants from Siberia likely crossed the Bering Strait with dogs in their company, and some experts suggest that use of sled dogs may have been critical to the success of the waves that entered North America roughly 12,000 years ago. Dogs were an important part of life for the Athabascan population in North America, and were their only domesticated animal. Dogs also carried much of the load in the migration of the Apache and Navajo tribes 1,400 years ago. Use of dogs as pack animals in these cultures often persisted after the introduction of the horse to North America.
The current consensus among biologists and archaeologists is that the dating of first domestication is indeterminate. There is conclusive evidence that dogs genetically diverged from their wolf ancestors at least 15,000 years ago, but some believe domestication to have occurred earlier. It is not known whether humans domesticated the wolf as such to initiate dog's divergence from its ancestors, or whether dog's evolutionary path had already taken a different course prior to domestication. The latter view has gained proponents, such as biologists Raymond and Lorna Coppinger; they theorize that some wolves gathered around the campsites of the paleolithical man to scavenge refuse, and that associated evolutionary pressure developed that favored those who were less frightened by, and keener in approaching, humans.
The bulk of the scientific evidence for the evolution of the domestic dog stems from archaeological findings and mitochondrial DNA studies. The divergence date of roughly 15000 years ago is based in part on archaeological evidence that demonstrates that the domestication of dogs occurred more than 15,000 years ago, and some genetic evidence indicates that the domestication of dogs from their wolf ancestors began in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. But there is a wide range of other, contradictory findings that make this issue controversial.
Archaeological evidence suggests that the latest dogs could have diverged from wolves was roughly 15000 years ago, although it is possible that they diverged much earlier. In 2008, a team of international scientists released findings from an excavation at Goyet Cave in Belgium declaring that a large, toothy canine existed 31,700 years ago and ate a diet of horse, musk ox and reindeer. Prior to this Belgium discovery, the earliest dog fossils were two large skulls from Russia and a mandible from Germany, that dated from roughly 14,000 years ago. Remains of smaller dogs from Natufian cave deposits in the Middle East, including the earliest burial of a human being with a domestic dog, have been dated to around 10,000 to 12,000 years ago. There is a great deal of archaeological evidence for dogs throughout Europe and Asia around this period and through the next two thousand years (roughly 8,000 to 10,000 years ago), with fossils uncovered in Germany, the French Alps, and Iraq, and cave paintings in Turkey.
DNA studies have provided a wider range of possible divergence dates, from 15,000 to 40,000 years ago, to as much as 100,000 to 140,000 years ago. This evidence depends on a number of assumptions that may be
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