A fish is any aquatic vertebrate animal that is typically ectothermic (or cold-blooded), covered with scales, and equipped with two sets of paired fins and several unpaired fins. Fish are abundant in the sea and in fresh water, with species being known from mountain streams (e.g., char and gudgeon) as well as in the deepest depths of the ocean (e.g., gulpers and anglerfish).
Food prepared from fish is also called fish, and it is an important food source for humans. They are harvested either from wild fisheries (see fishing) or farmed in much the same way as cattle or chickens (see aquaculture). They are also exploited by recreational fishers and fishkeepers, and are exhibited in public aquaria. Fish have had a role in many cultures through the ages, ranging from deities and religious symbols to the subjects of books and popular movies.
Diversity of fish
Main article: Diversity of fishThe term "fish" is most precisely used to describe any non-tetrapod craniate (i.e. an animal with a skull and in most cases a backbone) that has gills throughout life and has limbs, if any, in the shape of fins. Unlike groupings such as birds or mammals, fish are not a single clade but a paraphyletic collection of taxa, including hagfishes, lampreys, sharks and rays, ray-finned fishes, coelacanths, and lungfishes.
A typical fish is ectothermic, has a streamlined body that allows it to swim rapidly, extracts oxygen from the water using gills or an accessory breathing organ to enable it to breathe atmospheric oxygen, has two sets of paired fins, usually one or two (rarely three) dorsal fins, an anal fin, and a tail fin, has jaws, has skin that is usually covered with scales, and lays eggs that are fertilized internally or externally.
To each of these there are exceptions. Tuna, swordfish, and some species of sharks show some warm-blooded adaptations, and are able to raise their body temperature significantly above that of the ambient water surrounding them. Streamlining and swimming performance varies from highly streamlined and rapid swimmers which are able to reach 10–20 body-lengths per second (such as tuna, salmon, and jacks) through to slow but more maneuverable species such as eels and rays that reach no more than 0.5 body-lengths per second. Many groups of freshwater fish extract oxygen from the air as well as from the water using a variety of different structures. Lungfish have paired lungs similar to those of tetrapods, gouramis have a structure called the labyrinth organ that performs a similar function, while many catfish, such as Corydoras extract oxygen via the intestine or stomach. Body shape and the arrangement of the fins is highly variable, covering such seemingly un-fishlike forms as seahorses, pufferfish, anglerfish, and gulpers. Similarly, the surface of the skin may be naked (as in moray eels), or covered with scales of a variety of different types usually defined as placoid (typical of sharks and rays), cosmoid (fossil lungfishes and coelacanths), ganoid (various fossil fishes but also living gars and bichirs), cycloid, and ctenoid (these last two are found on most bony fish). There are even fishes that spend most of their time out of water. Mudskippers feed and interact with one another on mudflats and are only underwater when hiding in their burrows. The catfish Phreatobius cisternarum lives in underground, phreatic habitats, and a relative lives in waterlogged leaf litter.
Fish range in size from the 16 m (51 ft) whale shark to the 8 mm (just over ¼ of an inch) long stout infantfish.
Many types of aquatic animals commonly referred to as "fish" are not fish in the sense given above; examples include shellfish, cuttlefish, starfish, crayfish and jellyfish. In earlier times, even biologists did not make a distinction – sixteenth century natural historians classified also seals, whales, amphibians, crocodiles, even hippopotamuses, as well as a host of aquatic invertebrates, as fish. In some contexts, especially in aquaculture, the true fish are referred to as finfish (or fin fish ) to distinguish them from these other animals.
Classification
Fish are a paraphyletic group: that is, any clade containing all fish also contains the tetrapods, which are not fish. For this reason, groups such as the "Class Pisces" seen in older reference works are no longer used in formal classifications.
Fish are classified into the following major groups:
- Subclass Pteraspidomorphi (early jawless fish)
- Class Thelodonti
- Class Anaspida
- (unranked) Cephalaspidomorphi (early jawless fish)
- (unranked) Hyperoartia or Petromyzontida
- Petromyzontidae (lampreys)
- Class Galeaspida
- Class Pituriaspida
- Class Osteostraci
- Infraphylum Gnathostomata (jawed vertebrates)
- Class Placodermi (armoured fishes, extinct)
- Class Chondrichthyes (cartilaginous fish)
- Class Acanthodii (spiny sharks, extinct)
- Superclass Osteichthyes (bony fish)
- Class Actinopterygii (ray-finned fish)
- Subclass Chondrostei
- Order Acipenseriformes (sturgeons and paddlefishes)
- Order Polypteriformes (reedfishes and bichirs).
- Subclass Neopterygii
- Infraclass Holostei (gars and bowfins)
- Infraclass Teleostei (many orders of common fishes)
- Class Sarcopterygii (lobe-finned fish)
- Subclass Coelacanthimorpha (coelacanths)
- Subclass Dipnoi (lungfish)
Some palaeontologists consider that Conodonta are chordates, and so regard them as primitive fish. For a fuller treatment of classification, see the vertebrate article.
The various fish groups taken together account for more than half of the known vertebrates. There are almost 28,000 known extant species of fish, of which almost 27,000 are bony fish, with the remainder being about 970 sharks, rays, and chimeras and about 108 hagfishes and lampreys. A third of all of these species are contained within the nine largest families; from largest to smallest, these families are Cyprinidae, Gobiidae, Cichlidae, Characidae, Loricariidae, Balitoridae, Serranidae, Labridae, and Scorpaenidae. On the other hand, about 64 families are monotypic, containing only one species. It is predicted that the eventual number of total extant species will be at least 32,500.
Anatomy
Main article: Fish anatomyDigestive system
The advent of jaws allowed fish to eat a much wider variety of food, including plants and other organisms. In fish, food is ingested through the mouth and then broken down in the esophagus. When it enters the stomach, the food is further broken down and, in many fish, further processed in finger-like pouches called pyloric caeca. The pyloric caeca secrete digestive enzymes and absorb nutrients from the digested food. Organs such as the liver and pancreas add enzymes and various digestive chemicals as the food moves through the digestive tract. The intestine completes the process of digestion and nutrient absorption.
Respiratory system
Most fish exchange gases by using gills that are located on either side of the pharynx. Gills are made up of threadlike structures called filaments. Each filament contains a network of capillaries that allow a large surface area for the exchange of oxygen and carbon dioxide. Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gill filaments. The blood in the capillaries flows in the opposite direction to the water, causing counter current exchange. They then push the oxygen-poor water out through openings in the sides of the pharynx. Some fishes, like sharks and lampreys, possess multiple gill openings. However, most fishes have a single gill opening on each side of the body. This opening is hidden beneath a protective bony cover called an operculum.
Juvenile bichirs have external gills, a very primitive feature that they hold in common with larval amphibians.
Many fish can breathe air. The mechanisms for doing so are varied. The skin of anguillid eels may be used to absorb oxygen. The buccal cavity of the electric eel may be used to breathe air. Catfishes of the families Loricariidae, Callichthyidae, and Scoloplacidae are able to absorb air through their digestive tracts. Lungfish and bichirs have paired lungs similar to those of tetrapods and must rise to the surface of the water to gulp fresh air in through the mouth and pass spent air out through the gills. Gar and bowfin have a vascularised swim bladder that is used in the same way. Loaches, trahiras, and many catfish breathe by passing air through the gut. Mudskippers breathe by absorbing oxygen across the skin (similar to what frogs do). A number of fishes have evolved so-called accessory breathing organs that are used to extract oxygen from the air. Labyrinth fish (such as gouramis and bettas) have a labyrinth organ above the gills that performs this function. A few other fish have structures more or less resembling labyrinth organs in form and function, most notably snakeheads, pikeheads, and the Clariidae family of catfish
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